The invention relates to a fuel cell (1), a fuel cell stack (10) having at least two fuel cells (1), a fuel cell device having a fuel cell stack and a motor vehicle having a fuel cell device. In order to prevent a production-related mispositioning of a membrane electrode assembly (13) of the fuel cell (1) from causing obstructions in the operating media flow, it is provided according to the invention that at least one of the bipolar plates (2, 3) projects farther into an operating media line (4) than the other of the bipolar plates (2, 3).